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- Traits:SublittoralFringe + (The upper part of the sublittoral zone whi … The upper part of the sublittoral zone which is uncovered by the tide. On hard substrata, the zone is characterized by the kelps ''Laminaria digitata'' and ''Alaria esculenta''. The lower limit of this zone is marked by the upper limit of the truly sublittoral kelp ''Laminaria hyperborea'' (based on Lewis, 1964; Hiscock, 1996).a'' (based on Lewis, 1964; Hiscock, 1996).)