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- Traits:LowerLittoralFringe + (The bottom of the littoral fringe. In the … The bottom of the littoral fringe. In the UK, characterized by he ''Pelvetia/Porphyra'' belt with patchy ''Verrucaria maura'' and ''Fucus spiralis'' (on sheltered shores). ''Verrucaria mucosa'' present above the main barnacle population. May also include salt marsh species on shale/pebbles in shelter (Hiscock, 1996). shale/pebbles in shelter (Hiscock, 1996).)